A
Canada goose is a Canada goose is a Canada goose, right? Wrong –
not anymore.
With the 45th supplement of its official checklist published in
July 2004, the American Ornithologist’s Union (AOU) split
the large and varied species Branta canadensis into two species:
Branta Canadensis, the Canada goose, and Branta hutchinsii, the
cackling goose.
Many ornithologists and birders have long argued for some sort
of split among the large and varied group of Canada geese, which
before the split contained North America’s largest goose,
the giant Canada goose as well as one of North America’s smallest
— the cackling goose.
According to field guide author David Allen Sibley, the AOU has
been studying the Canada complex for a number of years with some
suggesting as many as four separate species. It appears the split
into two groups was based primarily on DNA studies.
The DNA showed that the entire Canada complex originated from
a common ancestor. However, it also showed considerable difference
between today’s smaller, more northerly nesters and their
larger more southerly cousins. Researchers believe that nesting
populations of Canadas became fragmented during the last Ice Age
and that the two groups have evolved separately since that time.
The two species break down this way: Branta canadensis (the large-bodied
group) includes seven subspecies; Branta c. canadensis, the Atlantic
Canada goose; Branta c. interior, the Hudson Bay Canada goose; Branta
c. maxima, the giant Canada goose; Branta c. moffitti, the Moffitt’s
or great basin Canada goose; Branta c. parvipes, the lesser Canada
goose; Branta c. occidentalis, the dusky Canada goose and Branta
c. fulva, the Vancouver’s Canada goose.
Branta hutchinsii, the cackling goose, includes four subspecies;
Branta h. hutchinsii, the Richardson’s cackling goose; Branta
h. taverneri, the Taverner’s cackling goose; Branta h. minima,
the cackling cackling goose and Branta h. leucopareia, the Aleutian
cackling goose.
For the most part it is easy to see the body and bill size differences
between the Canada goose and the cackling goose, and it’s
easy to hear the difference in voice (cackling has a higher-pitched
call) making the split look like a simple matter. But nature aficionados
know nature is never that tidy.
Between the extremes of large-bodied Canadas and small-bodied
cackling geese are a couple of intermediate species of relatively
the same size and bill dimensions. They are Branta c. parvipes the
lesser Canada goose and Branta h. taverneri the Taverner’s
cackling goose.
The lesser Canada is said to be lighter overall with a paler breast
but birders, ornithologists and biologists say there are many intermediate
individuals. In fact, in The Birds of North America Mowbray et al
listed all the Canadas on the North Slope of Alaska as taverneri/parvipes.
And while field studies suggested widespread interbreeding, DNA
testing showed little evidence of hybridization and placed the lesser
squarely in the Canada group and Taverner’s with the cackling
goose. Scientists say the similarity between the two species is
not caused by hybridization but rather by a process known as evolutionary
convergence – two species evolving towards each other.
Birders in the field confronted with distinguishing between Taverner’s
and lesser will have to rely on a combination of factors and may
still be left with a guess. The most notable differences are probably:
Taverner’s darker overall with darker breast, chunkier more
compact body with square-looking head and bill; lesser is lighter
with lighter breast, more slender body and rounded head.
Most birders in the East won’t be faced with that dilemma.
The most common cackling goose in the East is likely Branta h. hutchinsii
the Richardson’s. However, eight cackling geese believed to
be Branta h. minima, the cackling cackling goose were reported from
Mattamuskeet during this year’s Christmas Bird Count.
The Richardson’s nests in Arctic Canada and winters in south
Texas and Mexico.
Richardsons resemble miniature Canada geese. They are closer in
size to a mallard, however. Their backs are more silvery or frosted
than their much larger cousins and their heads are square and steep
with short, stubby bills.
Branta h. minima is primarily a western species nesting on the
northwest coast of Alaska and wintering in southern California.
They are even a bit smaller than Richardson’s and darker overall
with a dark breast, rounded head and short stubby bill.
The cackling goose was apparently proud of its new species designation
and showed up on a couple of east Tennessee Christmas counts as
well as the Balsam count and the Mattamuskeet count in North Carolina.
One individual was also recorded on Lake Julian in Asheville.
The new nomenclature is a bit quirky. Cackling goose is the common
name given to the new species. In the past it referred to the smallest
of the subspecies Branta c. minima. But, following scientific protocol,
the new species name Branta hutchinsii is derived from the first
named subspecies. This is how we are left with the redundant cackling
cackling goose for Branta h. minima.